西部太平洋に分布する分類群に重点を置いたフリソデウオ科 (アカマンボウ目) の分類学的レビュー
Jennifer M. Martin and Eric J. Hilton (2021). A taxonomic review of the family Trachipteridae (Acanthomorpha: Lampridiformes), with an emphasis on taxa distributed in the western Pacific Ocean. Zootaxa 5039 (3): 301–351.
https://doi.org/10.11646/zootaxa.5039.3.1
https://www.biotaxa.org/Zootaxa/article/view/zootaxa.5039.3.1
■Introduction
フリソデウオ科（リボンフィッシュ Ribbonfishes、ディールフィッシュ Dealfishes、およびその近縁種）は、地球全体に分布しており、3属に少なくとも10種が存在する（ユキフリソデウオ属 Zu Walters & Fitch 1960、フリソデウオ属 Desmodema Walters & Fitch 1960、およびサケガシラ属 Trachipterus Goüan 1770、図 1A）。これらの魚（図 1～3）は、細長く、極端に横方向に圧縮された体、大きな目、肋骨がなく、側線鱗に棘があること（Olney ら 1993）、大きく突き出た口（Ferry ら 2019）、成魚には腹鰭がないことが特徴である。また、個体発生中に劇的な形態変化が起こることでも知られている（Hubbs 1925）。フリソデウオ科の分類学と生物地理学に関する研究の多くは、地域的な記述（例：南西太平洋、Ogilby 1898、ニュージーランド、Hamilton 1916、地中海と北東大西洋、Palmer 1961、Lipej et al. 2018、Macali et al. 2020）、北西大西洋、Purchase 2017、東太平洋、Fitch 1964、Angulo & López-Sánchez 2017、タスマニア、Scott 1983、南アフリカ海域、Heemstra & Kannemeyer 1984、日本、Hayashi 2002、韓国、Ji et al. 2009、北太平洋、Savinykh & Baitalyuk 2011）に依存しており、形態学的データは限られており、分析も不完全である。これらの局所的なレビューは地域の生物多様性評価に役立つが、そのような記載では、広範囲に分布している、あるいは地球全体に分布しているとされる種の個体群レベルの違いが説明されない可能性がある。
姉妹群である oarfishes リュウグウノツカイ科 (Regalecidae、Olney ら、1993、図 1B) と同様に、フリソデウオ科は体が壊れやすい (つまり、収集中に損傷を受けることが多い) ため、コレクションにあまり含まれていない。フリソデウオ科の研究のほとんどは、調査された標本数、発育段階、および完全性によって制限されている。成魚に比べて、幼魚のフリソデウオ科は魚類コレクションでより一般的である。
これは、成魚の開放的な外海、多くの場合、深海の生息地と比較して、その個体数が多く、沿岸に近い生息地を占めていることによると考えられる。さらに、稚魚の標本の形態が独特なため、科レベルでは適切に識別できるが、より細かいレベルでは識別できない。フリソデウオ科の分類学レビューや識別キーでは、サケガシラ属とフリソデウオ属の成魚標本とユキフリソデウオ属の稚魚標本を比較し、種の記載をその標本に基づいて行うことは珍しくない (例: Walters & Fitch 1960、Palmer 1961、Fitch 1964、Ji et al. 2009)。比較をさらに複雑にしているのは、フリソデウオ科が個体発生中に受ける劇的な形態変化が、サイズと直接相関していないことである。フリソデウオ科魚類で稚魚の特徴が失われる時期は様々で、ユキフリソデウオ属では比較的遅い時期に発生することがあり、体長 800 mm までの標本では、長い背びれと腹びれの鰭条や、波型腹縁などの稚魚の特徴が保持される (Heemstra & Kannemeyer 1984; JMM、個人観察)。サケガシラ属の標本では、体長 600 mm まで、長い鰭条や斑点状の色素パターンなどの稚魚の特徴が保持される (JMM、個人観察)。Walters & Fitch (1960)、Palmer (1961)、Fitch (1964) による一般的な記載に基づいて、これらの研究における種の診断は、たとえば亜成魚のユキフリソデウオ属のみに基づいていた。種の記載が限られていることが、多くのコレクションで成魚の標本が誤って識別される理由の１つであると考えられる。Heemstra & Kannemeyer (1984) などの最近の研究では、コレクション内の成魚標本の希少性が認められているが、その診断は限定されており、南アフリカ海域で採取された Z. cristatus Bonelli 1820 の成魚標本１個体と Z. elongatus Heemstra & Kannemeyer 1984 の成魚標本２個体のデータのみが含まれている。
入手可能な資料の不足に加え、不完全で矛盾した特徴情報が、フリソデウオ科の分類上の混乱をさらに複雑にしている。例えば、Walters & Fitch (1960)、Palmer (1961)、Fitch (1964)、Scott (1983)、Heemstra & Kannemeyer (1984) はいずれも、サケガシラ属の診断的特徴として、あらゆる成長段階で鱗がないことを指摘しており、これがサケガシラ属をユキフリソデウオ属と区別するのに使われ、一部の著者によると、サケガシラ属をフリソデウオ属と区別するのにも使われている (Rosenblatt & Butler 1977)。しかし、本研究で調べたサケガシラ属の成体標本には鱗がある。したがって、鱗がないと推定されるのは、調査対象の標本の状態によるものであり、属の診断に使うことはできない。フリソデウオ科の属については、発育段階別に区分された世界規模のデータの統合を含む分類学上の改訂が明らかに必要である。これらの魚はコレクションでは比較的まれであるため、研究資料が限られているため、科の包括的なレビュー（地理と個体発生の観点から）が妨げられている。コレクションの資料が蓄積され、海洋の新しい地理的領域がサンプリングされるにつれて、特に成体段階の標本が検査に利用できる数が増える。以前の著者による研究における個体発生と地理的連続体の両方のギャップを埋めるために利用できる標本が増えているため、このレビューはタイムリーである。
フリソデウオ科の分類学を調べた地域的な改訂にもかかわらず、個体発生全体にわたる一連の形態学的特徴を統合したものはない。また、最近の分類学研究で西太平洋のリボンフィッシュの包括的なレビューに焦点を当てたものはない。この科の発育、生物地理、分類に関する文献の混乱により、世界中から集められたサケガシラ属の新しく収集された資料と博物館の標本が調査された。この研究の目的は、1) フリソデウオ科を改訂する、2) サケガシラ属、ユキフリソデウオ属、フリソデウオ属を改訂し、発育と生物地理に関する情報を組み込む、3) 西太平洋（日本、オーストラリア、ニュージーランド周辺の海域に重点を置く）のサケガシラ属、ユキフリソデウオ属、フリソデウオ属のアルファ分類について議論することである。
■References
Amaoka, K., Ama, G., Nakaya, M., Yabe, K., Masuda, K. & Sasaki, S. (1992) Larval and juvenile fishes collected from the northern Pacific Ocean by T/S Oshoro-Maru. Part I. Salmoniformes, Clupeiformes, Myctophiformes, Cyprinodontiformes, Lophiiformes, and Lampridiformes. Bulletin of the Faculty of Fisheries, Hokkaido University, 43, 1−23.
Angulo, A. & López-Sánchez, M.I. (2017) New records of lampriform fishes (Teleostei:Lampriformes) from the Pacific coast of lower Central America, with comments on the diversity, taxonomy, and distribution of the Lampriformes in the eastern Pacific Ocean. Zootaxa, 4236 (3), 573−591. https://doi.org/10.11646/zootaxa.4236.3.11
Backus, R.H., Mead, G.W., Haedrich, R.L. & Ebeling, A.W. (1965) The mesopelagic fishes collected during Cruise 17 of the R/V Chain, with a method for analyzing faunal transects. Bulletin of the Museum of Comparative Zoology, 134 (5), 139−157.
Bauchot, M.L. & Bianchi, G. (1984) First record of Desmodema polystictum (Ogilby, 1898) in North Indian Ocean. Cybium, 8 (4), 96−98.
Beltrán-León, B.S. & Herrera, R.R. (2000) Estadios tempranos de peces del Pacifico Colombiano. Tomos 1 & 2. Republica de Colombia, Ministerio de Agricultura y Desarrollo Rural, Instituto Nacional de Pesca Y Acuicultura (INPA), Buenaventura, Colombia, 726 pp.
Bleeker, P. (1868) Description et figure d’une nouvelle espèce de Trachypterus de l’île d’Amboine. Archives néerlandaises des sciences exactes et naturelles, 3, 279−280, pl. 12.
Bloch, M.E. & Schneider, J.G. (1801) Gen. 101. Trachypterus. In: Blochii, M.E. (Ed.), Systema Ichthyologiae Iconibus cx Ilustratum. Post obitum auctoris opus inchoatum absolvit, correxit, interpolavit Jo. Gottlob Schneider, Saxo. umtibus Austoris Impressum et Bibliopolio Sanderiano Commissum, Berolini [Berlin], pp. 480−483. https://doi.org/10.5962/bhl.title.5750
Bolin, R.L. (1933) New fish records from southern California. Copeia, 1, 35−36. https://doi.org/10.2307/1436190
Bonelli, F.A. (1820) Description d’une nouvelle espèce de poisson de la Méditerranée appartenant au genre Trachyptère avec des observations sur les caractères de ce même genre. Memorie della Reale Accademia delle Sciene di Torino, 24 (8129), 485−494, pl. 9.
Bounhiol, J.P. (1923) Une espèce nouvelle de Trachypterus Goüan: le Trachypterus Gavardi, Bounhiol. In: Bounhiol, J.P. & Gavard, A., Bulletin de l’Institut Océanographique, Monaco, 432, pp. 1−4.
Brünnich, M.T. (1788) Om den Islandske Fisk, Vogmeren. Gymnogaster arcticus. Kongelige Danske Videnskabernes Selskab, 3, 408−413.
Charter, S.R. & Moser, H.G. (1996) Trachipteridae. In: Moser, H.G. (Ed.), The early stages of fishes in the California current region. CalCOFI Atlas, 33, pp. 669−677.
Clark, H.W. (1938) Additional new fishes. The Templeton Crocker Expedition of 1934−35. No. 36. Proceedings of the California Academy of Sciences, Series 4, 22 (7), 179−185.
Clarke, F.E. (1881) Description of a new species of Trachypterus. Transactions of the New Zealand Institute, 13, 195−199.
Cocco, A. (1838) Osservazioni intorno taluni pesci del mare di Messina. Lettera. Il Faro Anno 6. 4 (15). Tip. M. Nobolo, Messina, 16 pp., 1 pl.
Costa, O.G. (1842) Fauna del regno di Napoli, ossia enumerazione di tutti gli animali che abitano le diverse regioni di questo regno e le acque che le bagnano, etc. Pesci. Fauna del regno di Napoli, 3 (pt 2), 1−148.
Cuvier, G. (1816) Le Règne Animal distribué d’après son organisation pour servir de base à l’histoire naturelle des animaux et d’introduction à l’anatomie comparée. Les reptiles, les poissons, les mollusques et les annélides. Vol.2. 1st Edition. xviii + 532 pp., pls. 9−10 in Vol. 4.
Deshmukh, A, Dragičevič, B., Paidi, C. & Muniraj. K. (2017) New record of Desmodema polystictum from northwestern Indian Ocean, Cybium, 41 (4), 379−381.
Dulčić, J. (2002) First record of scalloped ribbon fish, Zu cristatus (Pisces: Trachipteridae), eggs in the Adriatic Sea. Journal of Plankton Research, 24 (11), 1245−1246. https://doi.org/10.1093/plankt/24.11.1245
Emery, C. (1879) Le metamorfosi del Trachypterus taenia. Atti della Accademia Nazionale dei Lincei, 3 (3), 8.
Fahay, M.P. (2007) Lampridiformes. In: Fahay, M.P. (Ed.), Early stages of fishes in the western North Atlantic Ocean. Vol. I. North Atlantic Fisheries Organization, Dartmouth, Nova Scotia, pp. 824−840.
Ferry, L.A., Paig-Tran, E.W., Summers, A.P. & Liem, K.F. (2019) Extreme premaxillary protrusion in the king-of-the-salmon, Trachipterus altivelis. Journal of Morphology, 280 (12) 1865–1870. https://doi.org/10.1002/jmor.21071
Fitch, J.E. (1964) The ribbonfishes (family Trachipteridae) of the eastern Pacific Ocean, with a description of a new species. California Fish and Game, 50, 228−240.
Fitch, J.E. & Schultz, S.A. (1978) Some rare and unusual occurrences of fishes off California and Baja California. California Fish and Game, 64 (2), 74−92.
Fourmanoir, P. (1969) Contents stomacaux d’Alepisaurus (poissons) dans le Sud-Ouest Pacifique. Office de la Recherche Scientifique et Technique d’Outre-mer Séries Océanographie, 7 (4), 51−60.
Fricke, R., Eschmeyer, W.N. & van der Laan, R. (Eds.) (2020) Eschmeyer’s Catalog of Fishes: Genera, Species, References. Electronic version Available from; http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp (accessed 30 November 2020)
Garcia-Barcelona, S, Garcia-Cancela, R., Cayuela, M.S., Fernandez-Peralta, L., de Carlos, A., Baon, R., Macias, D. & Baez, J.C. (2016) Descripcion de dos ejemplares de Zu cristatus (Bonelli 1820) capturados accidentalmente con un palangre semipelagico en el Mediterraneo occidental. Arxius de Miscel-Iania Zoologica, 14, 91–98. [in Spanish] https://doi.org/10.32800/amz.2016.14.0091
Gmelin, J.F. (1789) Pisces in Caroli a Linné, Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Vol. 1. Part 3. impensis Georg. Emanuel. Beer, Lipsiae, pp. 1126−1516.
Goodrich, E.S. (1909) Part IX. Vertebrata Craniata (First Fascicle: Cyclostomes and Fishes). In: Lankester, R. (Ed.), A Treatise on Zoology. Adam and Charles Black, London, pp 1−518. https://doi.org/10.5962/bhl.title.13773
Goüan, A. (1770) Historia piscium, sistens ipsorum anatomen externam, internam, atque genera in classes et ordines redacta. Histoire des poissons, contenant la description anatomique de leurs parties externes et internes et le caractère des divers genres rangés par classes et par ordres. Amand König, Argentorati (Strasbourg), xviii + 252 (x2) pp., pl. IIV. [in Latin and French] https://doi.org/10.5962/bhl.title.6764
Günther, A. (1861) Trachypteridae. In: Günther, A. (Ed.), Catalogue of the fishes in the British Museum. Vol. 3. British Museum (Natural History), Department of Zoology, London, pp. 300−312.
Günther, A. (1887) Report on the deep-sea fishes collected by H. M. S. Challenger during the years 1873−76. Report on the Scientific Results of the Voyage of H. M. S. Challenger, 22 (57), i−lxv + 1−268, pls. 1−66. https://doi.org/10.5962/bhl.title.15693
Hamilton, H. (1916) Notes on the occurrence of the genus Trachipterus in New Zealand. Transactions and Proceedings of the Royal Society of New Zealand, 48, 370−382.
Harrisson, C.M.H. & Palmer, G. (1968) On the neotype of Radiicephalus elongatus Osório with remarks on its biology. Bulletin of the British Museum of Natural History, Zoology, 16, 187−211. https://doi.org/10.5962/p.144463
Hayashi, M. (2002) Trachipteridae. In: Nakabo, T. (Ed.), Fishes of Japan with pictorial keys to the species. English Edition. Tokai University Press, Tokyo, pp. 406−1489.
Heemstra, P.C. & Kannemeyer, S.X. (1984) The families Trachipteridae and Radiicephalidae (Pisces, Lampridiformes) and a new species of Zu from South Africa. Annals of the South African Museum, 94 (2), 13−39.
Herre, A.W. & Herald, E.S. (1950) Noteworthy additions to the Philippine fish fauna with descriptions of a new genus and species. Philippine Journal of Science, 79 (3), 309−340.
Hubbs, C.L. (1925) The metamorphosis of the Californian ribbon fish, Trachypterus rex-salmonorum. Papers of the Michigan Academy of Science, Arts and Letters, 5, 469−476.
Hutchins, J.B. (2001) Checklist of the fishes of Western Australia. Records of the Western Australian Museum, Supplement No. 63, 9−50. https://doi.org/10.18195/issn.0313-122x.63.2001.009-050
Hutton, F.W. (1873) Contributions to the ichthyology of New Zealand. Transactions of the New Zealand Institute, 1873, 264.
Hutton, F.W. (1876) Contributions to the ichthyology of New Zealand. Transactions of the New Zealand Institute, 1876, 214.
Ji, H.W., Yoon, S.C. & Kim, J.K. (2009) Taxonomic review of the family Trachipteridae (Lampridiformes) from Korea. Korean Journal of Ichthyology, 21 (4), 273−282. [in Korean]
Jordan, D.S. & Gilbert, C.H. (1882) Synopsis of the fishes of North America. Bulletin of the US National Museum, 16, 618−619. https://doi.org/10.5479/si.03629236.16.i
Jordan, D.S. & Gilbert, C.H. (1894) Description of a new species of ribbon fish Trachypterus rex-salmonorum, from San Francisco. Proceedings of the California Academy of Sciences, 2 (4), 144−146, pl. 9.
Jordan, D.S. & Snyder, J.O. (1901) Descriptions of nine new species of fishes contained in museums of Japan. Journal of the College of Science, Imperial University of Tokyo, 15 (2), 301−311, pls. 15−17.
Jordan, D.S. & Snyder, J.O. (1904) On a collection of fishes made by Mr. Alan Owston in the deep waters of Japan. Smithsonian Miscellaneous Collections, 45, 230−240, pls. 58−63.
King, J. E. & Ikehara, I. I. (1956) Some unusual fishes from the central Pacific.Pacific Science, 10, 17−24.
Kner, R. (1859) Ueber Trachypterus altivelis und Chaetodon truncatus, n. sp. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch- Naturwissenschaftliche Classe, 34, 437−445, pls. 1−2.
Leapley, W.T. (1953) First record of the ribbonfish, Trachipterus trachyurus, from the mainland of North America. Copeia, 4, 236. https://doi.org/10.2307/1440364
Lipej, L., Trkov, D. & Mavrič, B. (2018) Occurrence of ribbon fish (Trachipterus trachypterus) in Slovenian waters (northern Adriatic Sea). Annales. Series historia naturalis, 28 (2), 129–134. https://doi.org/10.19233/ASHN.2018.16
Lowe, R.T. (1852) An account of fishes discovered or observed in Madeira since the year 1842. Proceedings of the Zoological Society of London, 18 (5), 247. [1850]
Macali, A., Semenov, A., Paladini de Mendoza, F., Dinoi, A., Bergami, E. & Tiralongo, F. (2020) Relative Influence of Environmental Factors on Biodiversity and Behavioural Traits of a Rare Mesopelagic Fish, Trachipterus trachypterus (Gmelin, 1789), in a Continental Shelf Front of the Mediterranean Sea. Journal of Marine Science and Engineering, 8 (581), 1−13. https://doi.org/10.3390/jmse8080581
Martin, J.M. (2015) Phylogeny, ontogeny, and distribution of the ribbonfishes (Lampridiformes, Trachipteridae). Ph.D. Thesis. The College of William & Mary, Williamsburg, Virginia, 217 pp.
https://scholarworks.wm.edu/cgi/viewcontent.cgi?article=2489&context=etd
Masuda, H., Amaoka, A., Araga, C., Uyeno, T. & Yoshino, T. (1984) The fishes of the Japanese Archipelago. Tokai University Press, Tokyo, 437 pp.
McCoy, F. (1886) Natural history of Victoria. Prodromous of the zoology of Victoria. J. Ferres, Melbourne, 38 pp. https://doi.org/10.5962/bhl.title.4821
Meek, A. (1890) On the structure of Trachypterus arcticus (the modern ribbon-fish). Studies from the Museum of Zoology at University College Dundee, 1 (6), 1−24, pls. I + II.
Mincarone, M.M., Lima, A.T. & Soto, J.M.R. (2001) Sobre a ocorrência do peixe-fita Trachipterus jacksonensis (Ramsay, 1881) (Lampridiformes, Trachipteridae) na costa Basileria. Mare Magnum, 1 (2), 121−124. [in Portuguese]
Moore, J.A., Hartel, K.E., Craddock, J.E. & Galbraith, J.K. (2003) An annotated list of deepwater fishes from off the New England region, with new area records. Northeaster Naturalist, 10 (2), 159−248. https://doi.org/10.1656/1092-6194(2003)010[0159:AALODF]2.0.CO;2
Moritz, T., Stumer, D., Jakobsen, K. & Jakobsen, J. (2015) Observations on two live specimens of Trachipterus arcticus (Lampridiformes: Trachipteridae) from the Azores. Cybium, 39 (1), 78−80.
Mundy, B.C. (2005) Checklist of the fishes of the Hawaiian Archipelago. Bishop Museum Bulletins in Zoology, 6, 1−703.
Nelson, J.S., Crossman, E.J., Espinosa Pérez, Findley, L.T., Gilbert, C. R., Lea, R.N. & Williams, J.N. (2004) Common and scientific names of fishes from the United States, Canada, and Mexico. American Fisheries Society Special Publication 29. 6th Edition. American Fisheries Society, Bethesda, Maryland, 386 pp.
Nishimura, S. (1964) Additional information on the biology of the dealfish Trachipterus ishikawai Jordan & Snyder. Bulletin of Japan Sea Regional Fisheries Research Laboratory, 13, 127−129.
Norman, J.R. (1922) Two new fishes from New Britain and Japan. Annals and Magazine of Natural History, Series 9, 10 (56), 217−218. https://doi.org/10.1080/00222932308632764
Ogilby, J.D. (1898) On a Trachypterus from New South Wales. Proceedings of the Linnean Society of New South Wales, 3, 646−659. https://doi.org/10.5962/bhl.part.12736
Okiyama, M. (1988) An atlas of the early stages of fishes in Japan. Tokai University Press, Tokyo, 1154 pp.
Okiyama, M. (2014) An Atlas of Early Stage Fishes in Japan Second Edition. Vols. 1 & 2. Tokai University Press, Kanagawa, li + 976 pp., XX pls.
Olney, J.E. (1984) Lampridiformes: development and relationships. In: Moser, H.G., Richards, W.J., Cohen, D.M., Fahay, M.P., Kendall Jr., A.W. & Richardson, S.L. (Eds.), Ontogeny and systematics of fishes. American Society of Ichthyologists and Herpetologists Special Publication No. 1. Allen Press, Lawrence, Kansas, pp. 368−379.
Olney, J.E., Johnson, G.D. & Baldwin, C.C. (1993) Phylogeny of Lampridiform Fishes. Bulletin of Marine Science, 52 (1), 137−169.
Olney, J.E. & Naplin, A. (1980) Eggs of the scalloped ribbonfish, Zu cristatus (Pisces: Trachipteridae) in the western north Atlantic. Copeia, 1, 165−166. https://doi.org/10.2307/1444156
Olney, J.E. & Richards, W.J. (2006) Trachipteridae: dealfishes, ribbonfishes. In: Richards, W.J. (Ed.), Early stages of Atlantic Fishes. An identification guide for the western central north Atlantic. Vol. 1. Taylor & Francis, New York, pp. 1019−1026. https://doi.org/10.1201/9780203500217
Palmer, G. (1961) The dealfishes (Trachipteridae) of the Mediterranean and north-east Atlantic. Bulletin of the British Museum of Natural History, Zoology, 7, 335−351. https://doi.org/10.5962/p.314163
Pertseva-Ostroumova, T.A. & Rass, T.S. (1973) Ichthyoplankton of the southeastern Pacific Ocean. Transactions of the Institute of Oceanology Academy, Nauk, SSSR, 94, 7−70. [in Russian, English translation by the Department of the Secretary of State of Canada]
Philippi, R.A. (1874) Ueber eine neue Art Trachypterus aus dem chilenischen Meere. Archiv für Naturgeschichte, 40 (Pt. 1), 117−121, pl. 3.
Poey, F. (1861) Memorias sobra la historia natural de la Isla de Cuba, acompañadas de sumarios Latinos y extractos en Francés. Tomo 2. Imprinta de Barcina, La Habana, 3, 337−442, pls. 1−19. [in Spanish] https://doi.org/10.5962/bhl.title.2485
Purchase, C.F. (2017) First report of Dealfish, Trachipterus arcticus (Lampriformes: Trachipteridae), from Canadian waters. Canadian Field-Naturalist, 131 (4), 325–327. https://doi.org/10.22621/cfn.v131i4.1912
Quéro, J.C., Hureau, J.C., Karrer, C., Post, A. & Saldanha, L. (1990) Check-list of the fishes of the eastern tropical Atlantic. UNESCO, Lisbon, 1490 pp.
Quigley, D.T.G. & Henderson, G. (2014) First record of the scalloped ribbonfish Zu cristatus (Bonelli 1819) (Lampriformes: Trachipteridae) from N.W. European waters. The Glasgow Naturalist, 26 (1), 103−104.
Rafinesque-Schmaltz, C.S. (1810) Caratteri di alcuni nuovi generi e nuove specie di animali (principalmente di pesci) e piante della Sicilia, con varie osservazioni sopra i medisimi. Per le stampe di Sanfilippo, Palermo, 105 pp., 20 pls. [reprint, 1967, Asher-Amsterdam] https://doi.org/10.5962/bhl.title.104418
Ramsay, E.P. (1881) On a new species of Regalecus, from Port Jackson. Proceedings of the Linnean Society of New South Wales, 1881, pp. 631−633, pl. xx https://doi.org/10.5962/bhl.part.15897
Ranzani, C. (1818) Descrizione di un pesce, che appartiene ad un nuovo genere della famiglia dei Tenioidi di Cuvier. Opuscoli Scientifici, 133−137, pl. VI. [in Italian]
Rathnasuriya, M.I.G., Mateos-Rivera, A., Bandara, A.G.G.C., Skern-Mauritzen, R., Jayasinghe, R.P.P.K., Krakstad, J.O. & Dalpadado, J. (2019) DNA barcoding confirms the first record of a Desmodema polystictum (Ogilby, 1898) egg and all-time high adult catches in the Indian Ocean. Marine Biodiversity Records, 12 (22), 1−7. https://doi.org/10.1186/s41200-019-0180-7
Richards, W.J. (1990) List of Fishes of the Western Central Atlantic and the Status of Early Life Stage Information. NOAA Technical Memorandum NMFS-SEFC, 267, 1–88.
Roberts, C.D. (1993) Comparative morphology of spined scales and their phylogenetic significance in the Teleostei. Bulletin of Marine Science, 52 (1), 60−113.
Roberts, T.R. (2012) Systematics, biology and distribution of the species of the oceanic oarfish genus Regalecus. Mémoires du Muséum national d’Histoire naturelle. Tome 202. Muséum National d’Histoire Naturelle, Paris, 268 pp.
Rosenblatt, R.H. & Butler, J.L. (1977) The ribbonfish genus Desmodema, with the description of a new species (Pisces, Trachipteridae). Fishery Bulletin, 75 (4), 843−855.
Sabaj, M.H. (2020) Codes for Natural History Collections in Ichthyology and Herpetology. Copeia, 108 (3), 593–669. https://doi.org/10.1643/ASIHCODONS2020
Sanzo, L. (1918) Uova e larve di Trachypterus cristatus. R. Comitato Talassografico Italaliano Memoria, LXIV, 1−16. [in Italian]
Savinykh, V.F. & Baitalyuk, A.A. (2011) Taxonomic status of ribbonfishes of the genus Trachypterus (Trachipteridae) from the northern part of the Pacific Ocean. Journal of Ichthyology, 51 (8), 581−589. https://doi.org/10.1134/S0032945211040175
Scott, E.O.G. (1983) Observations on some Tasmanian fishes: Part XXIX. Papers and Proceedings of the Royal Society of Tasmania, 117, 167−202. https://doi.org/10.26749/rstpp.117.167
Scott, E.O.G. (1984) Observations on some Tasmanian fishes: Part XXX. Papers and Proceedings of the Royal Society of Tasmania, 118, 187−222. https://doi.org/10.26749/rstpp.118.187
Shinohara, G., Endo, K., Matsuura, K., Machida, Y. & Honda, H. (2001) Annotated checklist of deepwater fishes from Tosa Bay, Japan. In: Fujita, T.H., Saito, H. & Takeda, M. (Eds.), Deep-sea fauna and pollutants in Tosa Bay. National Museum of Nature and Science, Monographs No. 20, pp. 283−343.
Shinohara, G., Masanori, N., Ueda, Y., Kojima, S. & Matsuura, K. (2014) Annotated checklist of deep-sea fishes of the Sea of Japan. In: Fujita, T. (Ed.), Deep-sea fauna of the Sea of Japan. National Museum of Nature and Science Monographs, No. 44, 225−291.
Shirke, S.S., Ramachandran, S., Pradeep, H.P., Musaliyarakam, N., Devi, S.M. & Sinha, M.K. (2017) First record of the Taper-tail ribbonfish Zu elongatus Heemstra & Kannemeyer, 1984 from Indian EEZ. Fishtaxa, 2 (1), 43−47.
Smith, J.L.B. (1953) The Sea Fishes of Southern Africa. Revised enlarged edition. Central News Agency, Johannesburg, 564 pp., 107 pls.
Smith, J.L.B. (1956) A new dealfish from South Africa. Annals and Magazine of Natural History, Series 12, 9, 1 (02), 449−452, pl. 12. https://doi.org/10.1080/00222935608655839
Snyder, J.O. (1908) Description of Trachypterus seleniris, a new species of ribbon-fish from Monterey Bay, California. Proceedings of the Academy of Natural Sciences of Philadelphia, 60, 319–320.
Sparta, A. (1933) Trachypteridae e Regalecidae. In: Faune e flora del Golfo di Napoli: Uova, larvae e stadi giovanii de Teleostei. Pubblicazioni della Stazione Zoologica di Napoli, Monograph 38, pp. 266–279. [in Italian]
Swainson, W. (1839) On the natural history and classification of fishes, amphibians, & reptiles, or monocardian animals. Vol. II. Spottiswoode & Co., London, vi + 448 pp. https://doi.org/10.5962/bhl.title.62140
Tanaka, S. (1908) Notes on some Japanese fishes, with descriptions of fourteen new species. Journal of the College of Science, Imperial University of Tokyo, 23 (7), 1−54, pls. 1−4.
Tortonese, E. (1958) Cattura di Trachypterus cristatus Bon. e note sui Trachypteridae del mare Ligure. Doriana, II (89), 1−5. [in Italian]
Valenciennes, A. (1835) s.n. In: Cuvier, G. & Valenciennes, A., Histoire naturelle des poissons. Tome dixième. Suite du livre neuvième. Scombéroïdes. Livre dixième. De la famille des Teuthyes. Livre onzième. De la famille des Taenioïdes. Livre douzième. Des Athérines. Vol. 10. Chez F. G. Levrault, Paris, pp. i−xxiv + 1−482, 2 pp., pls. 280−306.
Van der Laan, R., Eschmeyer, W.N. & Fricke, R. (2014) Family-group names of Recent fishes. Zootaxa, 3882 (1), 1–230. https://doi.org/10.11646/zootaxa.3882.1.1
Walbaum, J.J. (1792) Petri Artedi sueci genera piscium. In quibus systema totum ichthyologiae proponitur cum classibus, ordinibus, generum characteribus, specierum differentiis, observationibus plurimis. Redactis speciebus 242 ad genera 52. Ichthyologiae Pars III. Ant. Ferdin. Rose, Grypeswaldiae [Greifswald], [viii] + 723 pp., pls. 1−3.
Walters, V. (1963) The trachipterid integument and a hypothesis on its hydrodynamic function. Copeia, 1963 (2), 260−270. https://doi.org/10.2307/1441341
Walters, V. & Fitch, J.E. (1960) The families and genera of the Lampridiform (Allotriognath) suborder Trachipteroidei. California Fish and Game, 46 (4), 441−451. https://doi.org/10.2307/1439670
Whitehead, P.J.P., Bauchot, M.L., Hureau, J.C., Nielsen, J. & Tortonese E. (1986) Fishes of the North-eastern Atlantic and Mediterranean. UNESCO, Paris, 491 pp. https://doi.org/10.2307/1444931
Wiley, E.O. & Johnson, G.D. (2010) A teleost classification based on monophyletic groups. In: Nelson, J.S., Schultze, H.-P. & Wilson, M.V.H. (Eds.), Origin and Phylogenetic Interrelationships of Teleosts. Verlag Dr. Friedrich Pfeil, München, pp. 123–182.
Young, S.S., Shen, S.C. & Chui, T.S. (1994). First record of trachipterid larvae with description of different stages and adult forms from Taiwan. Acta Zoologica Taiwanica, 5 (2), 85−93.
Zacharia, P.U. & Kannan, K. (2012) First record of polka-dot ribbonfish Desmodema polystictum (Pisces: Trachipteridae) from Indian waters. Marine Biodiversity Records, 5, 1−4. https://doi.org/10.1017/S1755267211001151
-----------------------------------------------------------------------------------------
Trachipteridae は「フリソデウオ科」ではなく、「サケガシラ科」にした方が筋道が通るのではなかろうか？
フリソデウオ科 Trachipteridae
■ユキフリソデウオ属 Zu
ユキフリソデウオ Z. cristatus (Bonelli, 1819)
■フリソデウオ属 Desmodema
フリソデウオ D. polystictum (Ogilby, 1898)
オキフリソデウオ D. lorum Rosenblatt and Butler, 1977
■サケガシラ属 Trachipterus
テンガイハタ T. trachypterus (Gmelin, 1789）
サケガシラ T. ishikawae Jordan and Snyder, 1901